Orchid Information - Family Orchidaceae Subfamily Orchidoideae Tribe Cranichideae Subtribe Goodyerinae

Goodyerinae Klotzsch (as ‘Goodyereae’) in Hayne, Getreue Darstell. Gew. (1846) 14, t. 24. Type genus: Goodyera R.Br.

Synonyms:

• Tribe Goodyerinae King & Pantl., Ann. Roy. Bot. Gard. Calcutta, 8 (1898) 273.
• Subtribe Physurinae Lindl., Gen: Sp. Orch. Pl., 442 (1840), [as Physuridae]. Type genus: Physurus L.C.Rich. ex Lindl., nom. illeg.
• Monandreae Neottiinae Physureae (Lindl.) Pfitzer, in Engler & Prantl, Die Natürl. Pflanzenf. II. 6. Orchidaceae (1889) n4. Type genus: Physurus L.C.Rich. ex Lindl., nom. illeg.
• Subtribe Pachyplectroninae Schltr. Notizbl. Bot. Gart. Mus. Berlin-Dahlem 9 (1926) 570. Type genus: Pachyplectron Schltr.
• Subtribe Cheirostylidinae Szlach., Folia Geobot. Phytotax. 26 (1991) 318. Type genus: Cheirosrylis Blume.

Perennial terrestrials, sometimes lithophytes and rarely epiphytic, herbaceous. Roots usually fleshy, hairy, from the nodes of the rhizome, rarely absent and replaced by rhizoids; velamen present or not. Rhizome creeping, short to elongate; internodes few to many, of equal length, rarely some abbreviated. Stem erect, terete, few- to many-leaved, rarely leafless. Leaves sessile, rarely reduced to sheaths and then without chlorophyll, or with a tubular leaf sheath and with a petiole-like base; leaf blade obliquely lanceolate to subcircular. lnflorescence terminal, erect, glabrous or with indument; peduncle usually with a few scale leaves; floral floral bracts lanceolate to ovate glabrous, glandular, or pubescent. Flowers resupinate or rarely not, glabrous or the sepals outside glandular or pubescent. Ovary cylindrical-fusiform, glabrous, glandular or pubescent. Sepals usually, rarely connate at the base. Median sepal concave, often forming a hood with the petals. Lateral sepals free or connate, often oblique at the base; lower half clasping the lip. Petals free or connate entire, sessile or with a claw, often hyaline, usually adnate to the dorsal sepal. Lip usually fleshy; attached to the base and usually also to the base of the column, entire, 2- or 3-lobed; hypochile concave, saccate or spurred, sometimes with a fleshy external flange; inside on both sides with or without 1 or few finger-shaped, stalked or sessile appendages, rarely all over the inside, sometimes with 2 keels; epichile sessile or clawed, the claw fringed or not, small to large, linear, ovate, moon-shaped or transversely 2-lobed; spur absent or present, entire or 2-lobed, tapering, cylindrical or club-shaped. Column short to long, sessile or with a stalk; anther usually erect, 2-loculate; pollinia 2, 2-partite or rarely entire, sectile, usually club-shaped, attenuated into short to elongate caudicles; clinandrium usually deeply concave; rostellum short to elongate, erect to porrect, often deltoid, slightly emarginate to deeply bifid, in some genera carrying a distinct, dorsal, stipes; stigma lobes free or connate under the base of the rostellum, sometimes subtended by 1 or 2 sterile appendages. Fruit usually cylindrical-fusiform, dehiscent.
(after Ormerod & Cribb 2003)

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